What's eating my titanopsis hugo-schlechteri?

Titanopsis hugo-schlechteri is a rugged South African mimicry succulent rarely targeted by pests indoors. Mealybugs are the primary threat, hiding between the tubercled leaves. Root mealybugs can go undetected until the plant wilts. Fungus gnats are occasional in overly moist compost. Its textured surface makes inspections harder than with smooth succulents.

White waxy fluff wedged between the rough tubercled leaf surfaces or at the soil line; plant looks dull and fails to firm up Use a fine brush dipped in isopropyl alcohol to reach between tubercles; drench with dilute neem or imidacloprid for root-zone bugs

Sudden wilting despite dry compost; powdery white coating on roots when unpotted Unpot and inspect roots; rinse off colonies; soak in dilute systemic insecticide; repot in fresh gritty mineral mix

Fine root damage from larvae in overly wet or peat-heavy compost Use near-soilless mineral compost; water infrequently; apply Bti drench if adults are visible